No tuning capacitor should ever show DC resistance (the only kind a DMM will measure)!
If that capacitor is shorted then it would completely explain the symptoms you've been discussing. Find that short and fix it.
Also if the grid lug of the first RF tube shows resistance to ground with the tube removed and nothing connected to it, then the socket is bad...Which means find and remove the conductive path or replace the socket.
Both of those problems will kill a whole lot of signal if left be.
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