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A couple of things:
The resistor that's smoking, if it tests fine then it's smoking because the short is on (or not far down stream of) the side of the resistor not connected to the high B+ rail....That resistors is the somewhat literal metaphorical smoking gun that points you to at least one killer short.
There were a few purposes to the relay. The main one is to delay the B+ until the tubes have warmed up to stable operation so when the set comes on you get a perfect picture and sound instantly without any concerning warmup instability. (Realistically you can delete the relay by shorting the contacts and the set will still work fine... At least for sets that work with the relay there) The other purpose is to protect most of the lytics from warmup surge.... Basically since the 5U4s are filament tubes they warmup almost instantly, but all the other tubes take ~11 sec.... During that time the B+ has no load from the signal/sweep tubes and skyrockets briefly this can be hard on the lytics and or require higher voltage lytics to cope.
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