The rectifier tube was probably for reverse voltage protection in self rectifying sets. After all if the rectifying part of the vibrator went bad and AC or reversed DC hit the lytics without a second tube rectifier to stop it the lytics would end with a bang, and an alarming (to the average consumer) amount of smoke. I don't know how the other parts would react to AC or reverse voltage, but I'm gonna guess it wouldn't exactly be good.
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