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FWIW, I would first suspect a hard B+ short-to-ground (possibly from the H.centering control or V.centering control to ground).
A hard short on the B+ line would throw a huge current draw on the 5U4 and its centertapped HV winding, and cause a large overcurrent through the resistor chain off the centertap, which includes R127B (it's the highest resistance in the chain, so it will run the hottest).
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