Replacing those tuner resistors didn't seem to have any effect so I went back to square one. It seemed to be that although I could feed a signal into the IF and get an image, I was having to use a really strong signal.
So I started swapping and wiggling tubes even though I had already tested them all.
Eureka! An image and sound

Only problem though is that it's inverted
I swapped out the video output tube while the set was running and briefly got a normal image as the tube warmed up. I'll investigate the video amp circuit more tomorrow.