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C1 and C2A form a voltage doubler for the B+. They are constantly charging and discharging. See how C1 is going directly to the AC line ? All the B+ power for the set is pumped through that cap. That's why having a low ESR is important.
If either cap is bad, they would not fully charge up and B+ would drop.
It's the speaker field coil and C3A that are more responsible for filtering out B+ ripple.
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