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It isn't that bad.
The 6K6 voltages can all be explained if R185 is connected to the -85V supply point instead of the +225V.
The 6AT6 could be something different. The grid pin 1 should be a bit negative, if it is near zero it could be passing too much current. That could cause pin 7, the plate, to be pulled too low. Check R180. Another test would to measure the voltage on pin 7 if you remove the 6AT6, it should go to about 225V. Check R179.
It would also be a good idea, one that is always a good idea to do early, is to measure the voltage supply points in the power supply. +225V, +150V, -3.5V -14V and -85V.
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