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A suggestion - from a dead-cold start, with your voltmeter on the 135V source (jct. of R185/R186), turn the set on. Since the 5U4s give full B+ almost instantly, is the 135V still low? Or is there a signifigant drop as the rest of the tubes begin conducting? This would tell you if it's tubes lugging the 135V down, or something else circuit-wise.
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